How to change breadcrumbs background color on crumb hover?
It appears that it uses the same color as "Caret Row" style .. and just automatically makes it lighter (darker the color the bigger difference will be):
This effect currently not disable.
Related issue: https://youtrack.jetbrains.com/issue/IDEA-139930
As the breadcrumb changed appearance in 2017.2, this can now also be configured better. Go to File > Settings > Editor > General > Breadcrumbs (Windows/Linux) or PhpStorm > Preferences > Editor > General > Breadcrumbs for macOS. There you can press "Manage colors" to get to the options pictured above.
https://www.jetbrains.com/help/phpstorm/navigating-with-breadcrumbs.html