I\'m getting this in my output Window:
System.Windows.Data Error: 4 : Cannot find source for binding with reference \'RelativeSource FindAncestor, Ancesto
I was also going to recommend Bea Stollnitz's article but Jonathan Allen got his post in while I was still typing this one. I also recommend the links in this blog entry.
In this particular case you can see that somewhere a ListBoxItem has a FindAncestor binding to an ItemsControl that is failing. That tells you right away there is a ListBoxItem somewhere that is either:
In addition, you know that someone, somewhere, is binding a ListBoxItem's VerticalContentAlignment property to FindAncestor.
Looking at the system themes (shipped with Expression Blend and also available through NET Reflector's BAMLViewer Add-in), we see this: